Know more

About cookies

What is a "cookie"?

A "cookie" is a piece of information, usually small and identified by a name, which may be sent to your browser by a website you are visiting. Your web browser will store it for a period of time, and send it back to the web server each time you log on again.

Different types of cookies are placed on the sites:

  • Cookies strictly necessary for the proper functioning of the site
  • Cookies deposited by third party sites to improve the interactivity of the site, to collect statistics

Learn more about cookies and how they work

The different types of cookies used on this site

Cookies strictly necessary for the site to function

These cookies allow the main services of the site to function optimally. You can technically block them using your browser settings but your experience on the site may be degraded.

Furthermore, you have the possibility of opposing the use of audience measurement tracers strictly necessary for the functioning and current administration of the website in the cookie management window accessible via the link located in the footer of the site.

Technical cookies

Name of the cookie


Shelf life

CAS and PHP session cookies

Login credentials, session security



Saving your cookie consent choices

12 months

Audience measurement cookies (AT Internet)

Name of the cookie


Shelf life


Trace the visitor's route in order to establish visit statistics.

13 months


Store the anonymous ID of the visitor who starts the first time he visits the site

13 months


Identify the numbers (unique identifiers of a site) seen by the visitor and store the visitor's identifiers.

13 months

About the AT Internet audience measurement tool :

AT Internet's audience measurement tool Analytics is deployed on this site in order to obtain information on visitors' navigation and to improve its use.

The French data protection authority (CNIL) has granted an exemption to AT Internet's Web Analytics cookie. This tool is thus exempt from the collection of the Internet user's consent with regard to the deposit of analytics cookies. However, you can refuse the deposit of these cookies via the cookie management panel.

Good to know:

  • The data collected are not cross-checked with other processing operations
  • The deposited cookie is only used to produce anonymous statistics
  • The cookie does not allow the user's navigation on other sites to be tracked.

Third party cookies to improve the interactivity of the site

This site relies on certain services provided by third parties which allow :

  • to offer interactive content;
  • improve usability and facilitate the sharing of content on social networks;
  • view videos and animated presentations directly on our website;
  • protect form entries from robots;
  • monitor the performance of the site.

These third parties will collect and use your browsing data for their own purposes.

How to accept or reject cookies

When you start browsing an eZpublish site, the appearance of the "cookies" banner allows you to accept or refuse all the cookies we use. This banner will be displayed as long as you have not made a choice, even if you are browsing on another page of the site.

You can change your choices at any time by clicking on the "Cookie Management" link.

You can manage these cookies in your browser. Here are the procedures to follow: Firefox; Chrome; Explorer; Safari; Opera

For more information about the cookies we use, you can contact INRAE's Data Protection Officer by email at or by post at :


24, chemin de Borde Rouge -Auzeville - CS52627 31326 Castanet Tolosan cedex - France

Last update: May 2021

Menu Logo Principal Institut Agro Montpellier LEPSE membre de University of Montpellier Labex AGRO Institut Carnot DigitAg

Home page

Koch Garance

Koch Garance
Garance Koch did a thesis on the development of tomato in response to water stress under the supervision of Nadia Bertin (PSH, Avignon) and Christine Granier (SPIC, LEPSE, Montpellier)

Thesis defended on 14.12.2018

Abstract :

As in other multicellular organs, leaf and fruit growth and development are characterised by the coordination of cell division and expansion, which are major growth processes. Cells in the pericarp of the fleshy fruit also undergo successive endocycles resulting in a significant increase in cell ploidy. There is a clear link between cell growth and endoreduplication, however, this is still poorly understood from a functional point of view. Cellular processes interact strongly during organ development and are linked to carbon and water flows in the plant. The objective of this thesis is to better understand the multi-scale control of leaf and fruit growth in tomato (Solanum lycopersicum Mill.) and the plasticity of their growth in response to soil water stress. The study focused on the cherry tomato genotype Solanum lycopersicum, cv.West Virginia 106 (WVa 106) which was grown under different irrigation conditions using automated systems developed for this study. The response to soil water deficit was studied at different scales of observation, (tissue, organ, whole plant) and at different stages of plant growth by adapting protocols used so far for determinate growth plants and single leaves. Two transgenic genotypes modified on a cell cycle regulation gene were also grown in order to vary the traits related to cell growth and better understand their links. The growth kinetics of the source and sink organs, i.e. the leaf and the fruit, at the cellular and tissue levels were also described. The results provided new information on the coordinations between the various processes studied and confirmed hypotheses already present in the literature. This work provided an original data set on the effects of water stress on the cellular processes (division, expansion, endoreduplication) involved in leaf and fruit growth in tomato, and enabled us to better understand their interactions on a larger scale, the plant as a whole. In terms of prospects, this data set could be used to develop a development model for fleshy fruit under optimal conditions and to test its genericity on another organ, the leaf. It opens up avenues for reflection on the modelling of plant plasticity in response to water stress.

Supervisors : Christine Granier & Nadia Bertin (PSH, Avignon, France)

Thesis project :

Multi-scale dynamic analysis of fruit and leaf growth in tomato in response to drought.

Publications :

Koch, G., Dambreville, A., Rolland, G., Dauzat, M., Chevalier, C., Frangne, N., Guedon, Y., Bertin, N., Baldazzi, V., Granier, C. (2016). . A multi-scale pipeline for reproducible analyses of tomato leaf expansion and its plasticity. Acta Horticulturae.

Bresson, J., Vasseur, F., Dauzat, M., Koch, G., Granier, C., Vile, D. (2015). . Quantifying spatial heterogeneity of chlorophyll fluorescence during plant growth and in response to water stress. Plant Methods, 11 (1), 23.